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Rome in 1074, and a decretal was issued that every layman who should receive the communion from the hands of a married priest should be excommunicated, and that every priest who married or lived in concubinage, should be deposed. The decree met with the most violent opposition in all countries; but Gregory succeeded in carrying it out with the greatest rigour; and though individual instances of married priests were still to be found in the 12th and 13th centuries, the C. of the Roman Catholic clergy was established, and has since continued both in theory and practice.

The violence thus done to human nature did not fail to avenge itself in those rude times. The licentiousness and corruption of the priests and monks became in many cases boundless, and it was in vain that strict individuals, as well as councils, strove against it. The immorality and debasement of the clergy became a reproach and by-word in the mouth of the people, and gave a powerful impulse to the religious movement that began in the 16th century. The leading Reformers declared against the C. of the clergy as unfounded in Scripture, and contrary to the natural ordinance of God, and Luther set the example of marrying. This was not without effect on the Roman Catholic clergy, and the question of the abolition of C. was raised at the council of Trent (1563). But the majority of voices decided that God would not withhold the gift of chastity from those that rightly prayed for it, and the rule of C. was thus finally and for ever imposed on the ministers of the Roman Catholic Church. Those who have only received the lower kinds of consecration may marry on resigning their office. For all grades above a subdeacon, a papal dispensation is necessary. A priest that marries incurs excommunication, and is incapable of any spiritual function. If a married man wishes to become a priest, he receives consecration only on condition that he separate from his wife, and that she of her free will consent to the separation and enter a religious order, or take the vow of chastity. The priests of the united Græco-Catholic congregations in Rome have received permission from the popes to continue in marriage, if entered into before consecration, but on condition of always living apart from their wives three days before they celebrate mass.

Notwithstanding these decisions, the contest against clerical C. has again and again been resumed, in recent times, both within and without the Roman Catholic Church. In fact, all attempts at innovation within the bosom of Catholicism, connect themselves with the attack on C., the abolition of which would deeply affect the constitution and position of that church. So far back as 1817, the Catholic Faculty of Tübingen expressed the opinion that compulsory C. was one of the chief causes of the want of Catholic ministers. In 1826, the Catholic clergy of Silesia put in petitions to the bishop for the abolition of C.; and unions were afterwards formed in Baden, Würtemberg, Bavaria, Silesia, and Rhenish Prussia, which, along with alterations in the doctrines and ritual of the Romish Church, combined attacks on the prohibition of marriage to the clergy. A work was also published, entitled The Introduction of Compulsory Celibacy among the Christian Priesthood, and its Consequences (Altenb. 1828, new ed. 1845), which excited great attention. At last the abolition of the law came to be discussed in the legislatures of Baden, Saxony, and other countries. The church claimed this subject as belonging exclusively to her jurisdiction, and not to that of the state; and in Würtemberg the clergy induced the government to suppress the anti-celibacy society; but this only made their opponents in the press the more zealous. In

France, also, the question, about 1829, was eagerly discussed. And in Spain, the Academy of Ecclesiastical Science took the subject into consideration in a meeting held in 1842; while the Portuguese Chambers had previously, in 1835, discussed it, though without result. The same took place in Brazil, about 1827.

During the commotions of 1848, the subject was again brought into prominence in Germany. The German Catholics (q. v.) had already abolished C.; and a general measure was called for in the Frankfort parliament, in the Prussian Assembly, and in the press. In Austria, also, voices were raised against it; but here the state took the side of the pope, who, in a bull of 1847, had added fresh stringency to the rule of C., and condemned its infringement. See BACHELOR.

CELL (Lat. cella, from celo, to conceal). The Latin word had nearly all the significations which we attach to the English one, and a good many besides which we have not borrowed. For example, the whole space within the walls of an ancient temple was called the cella. But the interior was frequently divided into several cellæ, in which case each C. took the name of the deity whose statue it contained, and was called the C. of Jupiter, Juno, Minerva, and the like. In these cases, the word approached to its general meaning, which, with the Romans as with us, was that of a store-room, or small apartment where objects of any kind were stowed away. In modern architecture, the term Vaulting C. signifies the hollow space between the principal ribs of a vaulted roof.

CE'LLÉ, or ZELL, a town of Hanover, on the left bank of the Aller, which at this point becomes navigable, 23 miles north-east of the city of Hanover. It is situated in the midst of a sandy plain, well built, and has a palace with a garden, in which Matilda, sister of George III., is buried. The inhabitants, about 10,000, are very industrious. The chief manufactures are linen, hosiery, tobacco, wafers, soap, &c. An active commerce is also carried on by the Aller, and by railway.

CELLI'NI, BENVENUTO, a celebrated Italian gold-worker, sculptor, founder, and medailleur, remarkable not only for his skill as an artist, but also for his checkered life, was born at Florence in the year 1500, and first displayed skill as a chaser and gold-worker. His autobiography is a remarkably curious and interesting work, presenting us with a complete picture of the author's life and character; his activity, his extraordinary weaknesses, the inpetuosity of his passions, the perilous circumstances in which his quarrelsome disposition placed him (for C. thought nothing of committing manslaughter in a moment of rage), and the ludicrous vanity and credulity which are never absent from him. The book is also of great value in a historico-social point of view, but does not impress us favourably in regard either to the personal or social morals of the time.

At an early period, having been banished from Florence in consequence of an 'affray,' C. went to Rome, where he was employed by many distinguished patrons of art, but afterwards was allowed to return to Florence. Another affray' compelled him to flee to Rome a second time, where he secured the favour of Clement VII. C., by his own account, was as great in arms as in art; he declares that it was himself who killed the Constable Bourbon and the Prince of Orange at the siege of Rome. His reckless conduct for some years compelled his constant shifting between Rome and Florence, Mantua, and Naples. In 1537 he went to the court of France, where he was very honourably received. Illness, however, induced him to return yet once more to

CELLINI-CELLS.

Rome, where he had the misfortune to be imprisoned on a charge of plundering the treasures in the castle of St Angelo during the siege of Rome. At length he was liberated, through the intercession of the Cardinal of Ferrara, for whom he executed, out of gratitude, a fine cup, and various other works. He now accompanied his deliverer to France, and entered the service of Francis I.; but having incurred the displeasure of the ruling favourite, Madame d'Estampes, he returned to Florence-not, however, until, as usual, he had settled some matters with his 'sword'-where, under the patronage of Cosmo de' Medici, he executed several fine works in metal and marble among them, the celebrated bronze group of Perseus with the Head of Medusa,' which now decorates the market-place in Florence. Among other preserved works of C., the splendid shield in Windsor Castle may be noticed. In his 58th year, he commenced writing his autobiography, and died in 1570 or 1572.

CELLS, in Physiology.-I. ANIMAL CELLS.-On examining, under a high magnifying power, any of the constituents of the animal body, we perceive that the smallest parts which appear to the naked eye as fibres, tubes, &c., are not ultimate elements in respect to form (morphotic elements), but that they contain and are built up of certain extremely minute particles, which differ in different organs, but always have a similar appearance in the same organs. By far the most important of these microscopic forms, which are known by histologists as 'simple elementary parts,' are the C., which not only form the starting-point of every animal and vegetable organism (the ovum in either kingdom of nature being simply a cell), but also either as C., or after having undergone certain modifications which will be presently describedmake up the tissues and organs of the perfect animal. Indeed, some of the lowest plants (red snow, gory dew), and of the simplest forms of animal life (GREGORINÆ, &c., q. v.), appear to consist of a single cell (see fig. 1).

[blocks in formation]

While in plants the elementary parts generally unite directly with one another, in animals they are usually combined by an interstitial substance, which may be either solid or fluid, and is always derived from the blood or general nutrient fluid. If this interstitial substance take a part in the formation of the C., it is called a cytoblastema or a blastema, from kutos, a cell or vesicle, and blastema, germsubstance; if it has nothing to do with their maintenance, it is called the matrix. The cytoblastema is usually fluid, as in the blood, chyle, &c.; while the matrix is solid, as in cartilage, bone, &c.

In every cell, we can distinguish, if we use sufficiently high magnifying powers, a membranous envelope, known as the cell-wall or membrane, and certain contents. The latter are fluid or gelatinous, and besides containing particles or granules, usually exhibit a peculiar rounded body, the nucleus; which, again, contains in its interior a fluid and a still smaller corpuscle, the nucleolus.

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forms may be mentioned: a, the polygonal, as in pavement epithelium, or the pigment of the eye; b, the conical or pyramidal, as in ciliated epithelium; c, the cylindrical, as in cylinder epithelium; d, the fusiform, or spindle-shaped, as in contractile fibre-C.; e, the squamous, as epidermic scales; and f, the caudate, polar, or stellate, as the C. in the gray nervous tissue (see fig. 2).

With regard to size, the largest animal C.-excepting the unicellular organisms-are the yolk-C. of the ova of birds and amphibia, while the blood-C. of certain animals may be taken as representing the smallest cells. Average C. range from 0·005 to 0-01 of a line in diameter.

The cell-membrane is usually transparent and colourless, mostly smooth, and so thin as to exhibit only a single contour, rarely of any measurable thickness. No traces of structure can be detected in it. The granular appearance which the membrane occasionally presents, is due to projections depending on granules lying on the inside; and it vanishes on the addition of water, which causes the cell to be distended by endosmosis. See OSMOTIC ACTION.

C. which contain only fluid are rare (fat-C., blood-C.); generally, besides fluid, they contain elementary granules and vesicles, and sometimes crystals. As a general rule, the number of these morphotic elements increases with the age of the cell; sometimes, however, this is not apparent, in consequence of their being grouped in a single mass around the nucleus.

The nucleus is usually spherical or lenticular, transparent, and either colourless or yellowish, and ranges from 0.002 to 0004 of a line in diameter. All nuclei are vesicles, as was originally maintained, in 1841, by Schwann (Microscopical Researches into the Accordance in the Structure and Growth of Animals and Plants, Sydenham Society's translation, 1847, p. 173), who must be regarded as the

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а

CELLS.

α

b

d

founder of the cell-theory in its relation to animal endogenous. Cartilage-C. afford a good example of tissues, and as has since been confirmed by Kölliker this process. The nucleus and the contents of each and other later observers. The contents of the parent cell undergo division into two parts, so that nucleus usually consist, the number of C. is successively doubled. This with the exception of process is exhibited in fig. 4, where a represents the nucleolus, of a limpid or slightly yellowish fluid, from which water and acetic acid precipitate granular matter. In general, only one nucleus exists in each cell, except when it is multiplying (a process which we shall presently explain); occasionally, however, we meet with several nuclei -four, ten, or even twenty (see fig. 3). The nucleolus

b

Fig. 3.

is

a, cells with a single nucleus; round, sharply defined,

b, a cell with two nuclei.

be almost immeasurable.

and often so small as to

Nucleoli are found in

As,

h

f

9

Fig. 4.

most nuclei so long as the latter are still young, and in many during their whole existence. however, nuclei exist in which no nucleolus can be the original cell; b, the same, beginning to divide; detected, we cannot regard the nucleolus as so essen-c, the same, shewing the complete division of the tial an element of the cell as the nucleus. Most nucleus; d, the same, with the halves of the nucleus commonly a nucleus contains only one nucleolus; separated, and the cavity of the cell subdivided; e, two are not unfrequently seen; more are rare. a continuation of the same process, with cleavage in a direction transverse to the first, so as to form a cluster of four C.; and f, g, h, the production of a longitudinal series of C., so as to produce filaments, by continuous cleavage in the same direction. The mode in which the multiplication of the nucleus takes place cannot be definitely made out in all cases, but when clear observation is possible, the nucleoli first divide into two, and then separate.

Our knowledge of the chemical composition of C. is very imperfect. That the cell-membrane is a protein substance (q. v.)-at all events in young C.-is obvious from its solubility in acetic acid and in dilute caustic alkalies; and the membrane of the nucleus seems to have a similar composition; while there are chemical reasons for believing that the nucleolus is composed of fat. In the contents of most C. we usually find such substances as occur in solution in the cytoblastema-viz., water, albumen, fat, extractive matters, and salts; and in the C. of secreting organs, as, for instance, the liver and kidneys, we find the special secretions of those glands; in the blood-C., we find hæmatocrystalline, &c.

There are two perfectly distinct ways in which C. can be generated: they may be developed independently of other C. in a plastic fluid (the cytoblastema); or they may be developed from preexisting C. by cell-multiplication, the existing C. either producing secondary C. within themselves, or multiplying by division. In both these latter kinds of cell-development, the nucleus seems to be the centre of development of the young cells.

In order that free or independent cell-development shall take place, we must have a cytoblastema containing protein substances (probably fibrin), fat, and certain salts (especially phosphates) in solution; and very possibly the presence of the particles of pre-existing C. may also be necessary, in which case free cell-development ceases to exist. The chyle and lymph corpuscles may be mentioned as examples of this mode of cell-formation. The steps of the process are not very clearly made out, but we know that the nuclei are first formed, and that the cell-membranes are developed around them. Free cell-development is far less common in man and the higher animals than cell-multiplication, and, we believe, never occurs in the vegetable kingdom. All pathological cell-formations-the C. in pus (q. v.), and in other morbid exudations-come, however, under this head.

The development of C. within other C. is of very common occurrence. An original or parent cell produces two or more secondary or daughter C., and the process of formation is said to be

A multiplication of C. by division has been proved to take place in the red blood-C. of the embryos of birds and mammals, and in the first colourless blood-C. of the tadpole, and very probably occurs extensively in many embryonic and adult tissues, in which a self-multiplication of C. is certain, but where no parent C. with secondary C. can be detected. În fig. 5 are shewn the blood-C. of the

8 8 8

Fig. 5.

Blood-corpuscles of the Chick, in the act of division.

chick dividing in this manner. In this and similar cases we have an elongation of the cell, and the single nucleus becomes divided into two; the cell then suffers constriction in the middle, which proceeds till it finally separates into two parts, each of which contains a nucleus. This variety of cellformation affords a good illustration of the doubt and difficulty connected with this class of investigations. It was altogether unknown to Schwann when he published his great work in 1839, and was first noticed and described by Remak in 1841, who, however, subsequently retracted his published view, and did not again advocate it till Kölliker confirmed his observation, and declared it to be correct.

No satisfactory theory has been propounded with the view of explaining the development of cells. Schwann compares the formation of C. with that of crystals, but it must be recollected that the molecular attraction concerned in the formation of

CELLS.

C. is so far peculiar, that-1. It never produces indiscriminately; but they have the power of geometrical solids, but even in the nucleus and taking up one constituent, and rejecting another, nucleolus determines a globular form; 2. That it and thus exhibit a selective faculty. aggregates not homogeneous, but chemically different substances; and 3. That the final result of its action-namely, the cell-is extremely limited in size, while a crystal may be of a comparatively indefinite magnitude.

same extent.

The cell having thus become filled from without, we have next to inquire into the changes which take place in the membrane and in the contents. As regards the former, the membranes of most C. not only become denser and more solid with The growth of C. requires some notice. Growth age, but they undergo changes in their chemical probably occurs in all Ĉ., although not in all to the constitution. Thus, in the horny tissues, the young It is most obvious in those which are C. are easily soluble in alkalies and acids, while formed directly round a nucleus, since in these the older C. of the same nature are scarcely affected membranes which at first closely invest the nucleus, by these re-agents; again, in cartilage-C., the in time become distended and enlarged, and merely membrane not only becomes firmer with age, and remain in contact with the nucleus at one point. thickens as ossification proceeds, but is changed Growth may take place either in surface or in thick- into a tissue yielding gelatine or glue on boiling, ness. The former is most commonly general-viz., which subsequently becomes impregnated with salts in all those cases where C. increase without alter- of lime (phosphate and carbonate). See BONE. ing their form; but is sometimes partial-viz., in The function of secretion is mainly carried on by those cases in which the cell deviates considerably changes in the contents of the cells. Thus, mucus from the primary globular form. The latter occurs is formed in the epithelial C. of the mucous memto a certain degree in all C., but branes, pepsin in those of the gastric glands, bile in some kinds to a far greater in the C. of the liver, and sepia in the C. of the extent than in others. In fig. 6, ink-bag of the cuttle-fish. In these cases, the C. two cartilage-C., magnified 350 do not separate mucus, pepsin, &c., from the blood, diameters, are shewn, in which the but merely the materials from which they elaborate walls are much thickened; in these substances. In other cases, as, for instance, in addition to their nucleus, each the C. of the kidney, the function of these minute contains a clear drop of fat. The organisms is not to manufacture new products, but nuclei and nucleoli also take part merely to separate certain substances (urea, uric Fig. 6. to a certain extent in the growth acid, &c.) from the blood, which, if not immediately of the cells. Schwann gives the removed from the general circulation, would speedily following general explanation of the process of accumulate, and act as a deadly poison. That these growth. He considers that the molecules of the C. merely separate the urea from the blood, and cell-membrane exert an attractive influence on the do not form it in their interior, is proved by the fluid which surrounds them, and deposit its newly fact that, if the kidneys of an animal are extirformed particles amongst themselves. If the depo-pated, the urea and other urinary constituents may sition take place between the molecules already speedily be found in large quantity in the blood. present in the substance of the membrane, the cell becomes distended; if it take place only in one or more definite directions, the membrane becomes thickened.

Having now traced the cell to the period of its full growth, we are prepared to consider the processes which occur in the interior of this minute organic structure, or, in other words, the physiology of cells. To enter satisfactorily into this subject, we ought to have an exact knowledge of the chemical composition of the contents of different cells. All that we know of the contents of C. generally is, as we have already stated, that they usually consist of a moderately concentrated solution of protein matters, with alkaline and earthy salts, and dissolved or suspended fat-particles; and that besides these ingredients many C. contain either a great preponderance of one of these constituents, to the almost entire exclusion of others, or are found to contain altogether new substances. Thus, there are C. with much protein matters, as the nerve-C., and with much fat, like the fat-C.; while there are other C. which specially contain hæmatine (the red colouring matter of the blood), pigment, biliary and urinary constituents, mucus, milk, sugar, &c.

The main cell-processes occurring in these variously constituted C. are absorption, secretion, and excretion. These depend principally, if not entirely, upon chemical and physical laws, and are to a great extent amenable to micro-chemical observation.

Absorption, or the appropriation of matters from without, is most manifest in those C. which at first have little or no contents save the nucleus. Although endosmose must be taken into account as a condition of absorption, C. must not be regarded merely as vesicles provided with indifferent porous membranes; for the filling of C. does not take place by their admitting every kind of matter

Excretion takes place by the bursting or solution of the distended secreting cell, usually into the duct of a secreting gland. The reader who desires further information on the functions of the C. in relation to secretion and excretion, is especially referred to an admirable memoir by Professor Goodsir, On Secreting Structures,' published in John and Harry D. S. Goodsir's Anatomical and Pathological Researches, 1845.

In conclusion, we must notice the metamorphoses of cells. The ovum itself is, as we have already mentioned, merely a nucleated cell; after impreg nation, a number of secondary C. are formed within it, by a process of cleavage or segmentation. See articles GENERATION and OVUM. Some of the C. which occur in the ovum in its early stages soon coalesce with others to form the higher elementary parts, which we shall shortly enumerate; others, without entering into combinations, more or less change their previous nature, as the horny plates of the epidermis and nails; while others, again, undergo no change of form throughout the period of their existence.

The permanent C. are arranged by Kölliker (Manual of Human Histology, translated by Busk and Huxley, 1853, vol. i. p. 47) under the following heads:

1. True Cells, which have in no essential respect altered their cellular character. These occur in the epidermis and the epithelium; in the blood, chyle, and lymph; in the glandular secretions, in the fatty tissue, in the gray nervous substance, in the glands (liver, spleen, &c.), and the cartilages. Their varieties of form and contents have been already noticed. Regarding their modes of occurrence, some are either isolated in fluids or in solid tissues; others are united by apposition, without any intervening structure, into a cellular parenchyma;

CELLS-CELLULAR TISSUE.

while others, again, are conjoined by an intercellular substance of some kind.

2. Metamorphosed Cells. To these belong-The horny scales: flattened, polygonal, or fusiform; their membrane being fused into one mass with their contents. They occur in the epidermis, the laminated pavement epithelium, and the hair and nails. The contractile fibre-C.: fusiform, slightly flattened, considerably elongated C., whose membrane, with its soft, solid contents, is changed into a contractile substance. They occur in the smooth or involuntary muscles. The tubules of the crystalline lens of the eye: very elongated C., with viscid, albuminous contents. The prisms of the enamel of the teeth: greatly elongated, prismatic, and strongly calcified cells. The bone-C.: thickened C. (with canaliculi, or minute branching canals) which have coalesced with the matrix of the bones. The transversely striated muscular C.: large polygonal C. whose contents have become metamorphosed into a transversely striated or striped tissue, such as is found in voluntary muscular fibre. From these C. are formed all the different fibres, net-works, membranes, tubes, &c.; in short, all the higher elementary parts of which the animal body is composed.

For further information on C. and cell-development, the reader is referred, in addition to the works quoted in this article, to Leydig, Lehrbuch der Histologie des Menschen und der Thiere, 1857; and to Frey, Histologie und Histochemie des Menschen, 1859; while he will find full details on morbid cell-development (the development and growth of C. in tubercle, cancer, and other morbid deposits) in Vogel's Pathological Anatomy of the Human Body, translated by Day, 1847; and in Wedl's Pathological Histology, translated (for the Sydenham Society) by Busk, 1855.

II. VEGETABLE CELLS.-In the vegetable, as in the animal kingdom, the primary form of the cell is that of a sphere. There are, however, interfering influences, which usually alter or modify the primary form, of which the most important are, (1.) Special directions assumed in the development, in obedience to a law regulating the structure of the tissue in which the cell occurs; and (2.) Obstructions to the expansion of the cell in certain directions from the pressure of surrounding cells.

The most common forms referrible to the law of development are, (1.) The spherical or fundamental form; (2.) The cylindrical, in which there is a tendency to elongation in the direction of a vertical axis; and (3.) The tubular, in which there is an excess of development in the direction of the two

transverse axes.

The secondary modifications of these forms are numerous. Thus, in lax tissues, the spherical form may become an irregular spheroid, running out into lobed, and even stellate forms, as may be seen in the pith of rushes and the stems of various aquatic plants. Again, in seeds, the hard part of fruits, &c., the mutual pressure of the C. converts the spherical into polyhedral forms, of which the dodecahedron giving a hexagonal section, and arising from equal pressure in all directions-is the most common, although cubic and many other forms occasionally

Both the cell-wall and the contents differ from the corresponding parts in animal cells. In all young C. the wall is membranous, freely permeable by water, elastic, and flexible. In many cases it retains these properties, whilst in others it becomes much modified, as the cell grows older. It consists mainly of CELLULOSE (q. v.). As the vital and chemical phenomena exhibited by plants depend primarily upon operations in the interior of the cell, the careful study of the cell contents is of the highest importance. Of these contents, the most important are the primordial utricle, with the protoplasm, the nucleus, chlorophyll corpuscles, and starch granules. The primordial utricle is a layer of substance of mucilaginous consistence (coloured yellow by iodine), lining the entire wall of the young cell, but often disappearing at a comparatively early period. The protoplasm is a tough mucilaginous and frequently granular fluid, which fills up the space in the interior of the cell not occupied by the nucleus. The nucleus or cytoblast is a globular or lenticular body, identical in its character with the substance of the primordial utricle, and occurring in the protoplasm of most young cells. Little is known with certainty regarding the chlorophyll corpuscles, except that, under the influence of solar light, green colouring matter is developed from them. Of the starch granules, which are very commonly found in the cell contents, we need not speak, as they are sufficiently described in the article STARCH.

In addition to the above organised structures, we must mention as frequent constituents of the cellcontents, fluid colouring matters, essential and fixed oils, resins, sugar, dextrine, gum, alkaloids, and mineral or organic salts, which are not unfrequently found in a crystalline form, when they are termed raphides.

There are two modes of cell-development in the vegetable kingdom-viz. (1.) Cell-division, where two or more new cells fill the cavity of the parent cell, and adhere to its membranes, appearing to divide it into compartments; and (2.) Free cell-formation—not to be confounded with a process of the same name which is supposed to occur in the animal kingdom-in which the whole or part of the cell-contents become detached from the cell-wall and resolved into new loose C., which ultimately escape from the parent cell. The former mode universally occurs in the formation of the C. by which growth is effected; the latter occurs only in the production of C. connected with reproduction. For further information, we must refer the reader to Von Mohl's Principles of the Anatomy and Physiology of the Vegetable Cell, translated by Henfrey, London, 1852.

CE'LLULAR TI'SSUE. This is the old term for a widely diffused animal texture, which has also received the names of areolar, reticular, filamentous, and connective tissue. If we make a cut through the skin, and proceed to raise it, we see that it is loosely connected with the subjacent parts by a soft, filamentous, elastic substance, which, when free from fat, has a white fleecy aspect. This is the tissue in question. It is also found underneath the serous and mucous membranes which are spread over internal surfaces, and serves to attach these membranes to the parts which they line. We likewise find it The magnitude of the vegetable C. is very varied. lying between the muscles, the blood-vessels, nerves, In flax, the liber-cells have been found, or even &c., occupying the interspaces between the different of an inch in length, and the cylindrical C. of organs, and often investing each of them with a some of the Confervæ are more than an inch long-special sheath. While it thus connects and insulates although their transverse diameter is very minutewhilst, on the other hand, the spores of Fungi are C. of a diameter of of an inch. The average diameter of the C. in the parenchymatous tissues is about of an inch.

occur.

entire organs, it at the same time performs a similar function in regard to the minute parts of which each organ is made up. Thus, for instance, in muscular tissue, it enters between the fibres of the muscle, uniting them into bundles; and similarly, it enters

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